Zealandia pustulata

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Family Polypodiaceae

Taxonomy: Zealandia pustulata (G.Forst.) Testo & A.R.Field

Synonyms: 13 (APC 2026-07-30)

Microsorium diversifolium Copel. (taxonomic synonym)
Microsorium pustulatum Copel. (nomenclatural synonym)
Microsorum diversifolium (Willd.) Copel. (taxonomic synonym)
Microsorum pustulatum (G.Forst.) Copel. (nomenclatural synonym)
Phymatodes diversifolia (Willd.) Pic.Serm. (taxonomic synonym)
Phymatodes diversifolium Pic.Serm. (taxonomic synonym)
Phymatodes pustulata (G.Forst.) C.Presl (nomenclatural synonym)
Phymatosorus diversifolius (Willd.) Pic.Serm. (taxonomic synonym)
Phymatosorus pustulatus (G.Forst.) Large, Braggins & P.S.Green (nomenclatural synonym)
Pleopeltis diversifolia (Willd.) Melvaine (taxonomic synonym)
Pleopeltis pustulata (G.Forst.) T.Moore (nomenclatural synonym)
Polypodium diversifolium Willd. (taxonomic synonym)
Polypodium pustulatum G.Forst. (nomenclatural synonym)

Misapplied names: 3 (APC 2026-07-30)

Polypodium phymatodes L. (misapplied)
Polypodium scandens G.Forst. (misapplied)
Polypodium scandens G.Forst. (pro parte misapplied)

Distribution (APC): Qld, NSW, LHI, NI, ACT, Vic, Tas

Profiles: APC, Flora of Australia, ALA, iNaturalist, NSW Flora, Vic Flora, ATRP

Traits

Top source datasets: ABRS_2022 (4), ABRS_2023 (8), Mokany_2015 (4), NHNSW_2022 (1), NHNSW_2023 (7), Smith_1996 (216), Wenk_2022 (2), Wenk_2023 (1), White_2020 (38), Williams_2011 (11). See below for a full list of sources.
Summary: AusTraits contains 52 traits from 292 records in 10 datasets.

Traits by category (click to expand). For categorical traits, the values are listed. For numerical traits, the mean and range are shown.

fire response

resprouting_capacity (categorical): fire_killed (2) [sources: White_2020]

leaf morphology

leaf_area (numerical): 3142.94 mm2 (573-5340.71 mm2) [sources: Mokany_2015; Smith_1996]
leaf_length (numerical): 251.88 mm (72-600 mm) [sources: Smith_1996; NHNSW_2023]
leaf_width (numerical): 19.49 mm (6-33 mm) [sources: Smith_1996; ABRS_2023]
leaf_dry_mass (numerical): 205.66 mg (62.5-283 mg) [sources: Smith_1996]
leaf_mass_per_area (numerical): 79.52 g/m2 (76.48-83.8 g/m2) [sources: Smith_1996]
leaf_dry_matter_content (numerical): 0.27 g/g (0.21-0.31 g/g) [sources: Smith_1996]
leaf_water_content_per_dry_mass (numerical): 2.7 g/g (2.18-3.74 g/g) [sources: Smith_1996]
leaf_margin (categorical): entire (2) [sources: ABRS_2023]
leaf_lobation (categorical): lobed (2) [sources: ABRS_2023; NHNSW_2023]
leaf_compoundness (categorical): simple (2) [sources: ABRS_2023; NHNSW_2023]
leaf_inclination_angle (numerical): 38.5 deg (15-90 deg) [sources: Smith_1996]
plant_photosynthetic_organ (categorical): leaf (2) [sources: ABRS_2023; NHNSW_2023]

life history

photosynthetic_pathway (categorical): c3 (1); cam (2) [sources: White_2020; Williams_2011]
plant_growth_form (categorical): climber (1); fern (2) [sources: ABRS_2022; Wenk_2022; Williams_2011]
life_form (categorical): chamaephyte (1) [sources: Williams_2011]
life_history (categorical): perennial (1) [sources: Wenk_2023]
plant_growth_substrate (categorical): epiphyte (4); lithophyte (1); terrestrial (1) [sources: ABRS_2022; NHNSW_2022; White_2020]
woodiness_detailed (categorical): herbaceous (2) [sources: ABRS_2022; Wenk_2022]
stem_growth_habit (categorical): climbing (1); creeping (1) [sources: ABRS_2022]
nitrogen_fixing (categorical): non_nitrogen_fixer (3) [sources: White_2020; Williams_2011]
root_structure (categorical): non_mycorrhizal (1) [sources: Williams_2011]
parasitic (categorical): not_parasitic (2) [sources: White_2020]
pollination_syndrome (categorical): abiotic (2) [sources: White_2020]
pollination_system (categorical): abiotic (1) [sources: Williams_2011]
dispersal_syndrome (categorical): anemochory (3) [sources: White_2020; Williams_2011]
dispersers (categorical): wind (2) [sources: White_2020]
seedbank_longevity_class (categorical): transient (2) [sources: White_2020]
vegetative_reproduction_ability (categorical): vegetative (2) [sources: White_2020]
clonal_spread_mechanism (categorical): rhizome (1) [sources: Williams_2011]
storage_organ (categorical): rhizome (1); rhizome_fleshy (1) [sources: ABRS_2023; NHNSW_2023]

nutrient and stable isotope

leaf_N_per_dry_mass (numerical): 9.9 mg/g (7.36-12.23 mg/g) [sources: Smith_1996]
leaf_P_per_dry_mass (numerical): 0.63 mg/g (0.46-0.84 mg/g) [sources: Smith_1996]

seed morphology

seed_dry_mass (numerical): 0 mg (0-0 mg) [sources: Williams_2011]

whole plant morphology

plant_height (numerical): 0.3 m (0.1-0.5 m) [sources: Mokany_2015]
plant_physical_defence_structures (categorical): absent (2) [sources: White_2020]

Sources

dataset_id	n_traits	n_records	reference
Smith_1996	17	216	B. Smith. “Community-level convergence and community structure of temperate Nothofagus forests”. PhD. University of Otago, Dunedin, New Zealand, 1996. url: http://web.nateko.lu.se/personal/benjamin.smith/thesis/thesis.html.
White_2020	19	38	M. White, S. Sinclair, and D. Frood. Victorian Vital Attributes Database. database. Department of Environment, Land, Water and Planning, Victoria, 2020.
Williams_2011	11	11	R. P. Duncan, S. E. Clemants, R. T. Corlett, A. K. Hahs, M. A. McCarthy, M. J. McDonnell, M. W. Schwartz, K. Thompson, P. A. Vesk, and N. S. G. Williams. “Plant traits and extinction in urban areas: a meta-analysis of 11 cities”. Global Ecology and Biogeography 20.4 (2011), pp. 509-519. doi: 10.1111/j.1466-8238.2010.00633.x.
ABRS_2023	6	8	Australian Biological Resources Study. Flora of Australia, Australian Biological Resources Study, Canberra.. accessed on 14 September, 2022.. 2023. url: http://www.ausflora.org.au.
NHNSW_2023	5	7	N. H. of New South Wales. The Plant Information Network System of The Royal Botanic Gardens and Domain Trust Version 2.0.. accessed on 6 May, 2022.. 2023. url: https://plantnet.rbgsyd.nsw.gov.au/.
ABRS_2022	4	4	Australian Biological Resources Study. Flora of Australia, Australian Biological Resources Study, Canberra.. accessed on 14 September, 2022.. 2022. url: http://www.ausflora.org.au.
Mokany_2015	2	4	K. Mokany, J. J. Thomson, A. J. J. Lynch, G. J. Jordan, and S. Ferrier. “Linking changes in community composition and function under climate change”. Ecological Applications 25.8 (2015), pp. 2132-2141. doi: 10.1890/14-2384.1.
Wenk_2022	2	2	E. Wenk. “Unpublished data: Australian plant growth forms dataset”. 2022.
NHNSW_2022	1	1	N. H. of New South Wales. The Plant Information Network System of The Royal Botanic Gardens and Domain Trust Version 2.0.. accessed on 6 May, 2022.. 2022. url: https://plantnet.rbgsyd.nsw.gov.au/.
Wenk_2023	1	1	E. Wenk. “Unpublished data: Australian plant life history dataset, University of New South Wales, Sydney”. 2023.